Pupulin, F. 2001. New taxa in Costa Rican Lepanthes (Orchidaceae). Harvard Pap. Bot. 6(1): 289-294.

Abstract
Two new orchid species and one subspecies are described and illustrated in the genus Lepanthes from Costa Rica: Lepanthes rafaeliana, L. spadarii, and L. johnsonii subsp. costaricensis. Ecological and taxonomic notes are provided for each taxon.

Keywords
Costa Rica, Lepanthes, Orchidaceae, taxonomy.

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